E3 SUMO-protein transferase PIAS2 zinc finger, MIZ-type containing 4 DIP ARIP3 PIASX DAB2-interacting protein msx-interacting zinc finger protein androgen receptor-interacting protein 3 E3 SUMO-protein ligase PIAS2 protein inhibitor of activated STAT X MIZ1 SIZ2 ZMIZ4