hVps32-3 chromatin-modifying protein 4c Shax3 charged multivesicular body protein 4c chromatin modifying protein 4C SNF7 homolog associated with Alix 3 vacuolar protein sorting-associated protein 32-3 SNF7-3 vps32-3 Snf7 homologue associated with Alix 3 hSnf7-3 VPS32C