Ohishi Y, Kinoshita R, Marubashi S, Ishida M, Fukuda M. The BLOC-3 subunit HPS4 is required for activation of Rab32/38 GTPases in melanogenesis, but its Rab9 activity is dispensable for melanogenesis. Journal of Biological Chemistry. 2019 Apr;294(17):6912–22. doi: 10.1074/jbc.ra119.007345.